Professor David Ellsworth

Professor Ellsworth is recognised as a "highly cited" researcher in the fields of ecology and environmental science, with a particular focus on the response of forests to climate change.

He served on the ARC "Excellence in Research for Australia" Biological Sciences committee (2010) as an expert in ecology and forestry. He is senior scientific advisor for the Eucalyptus Free-Air CO2 Enrichment (EucFACE) experiment.

He brings over 15 years experience in teaching and research at Western Sydney University and previously the University of Michigan, Duke University and Brookhaven National Laboratory in the USA. He has also been involved in many national and international conferences.

Communication of research on climate change to the wider community is one of Professor Ellsworth's passions. In addition to over 85 research articles in the international literature, he has written articles or been interviewed for Sydney's Sun-Herald, Reuters News Agency, the Washington Post, The New York Times and New York's Newsday, and National Geographic Magazine. He has appeared on news and science programs for Australian television.

David is currently principal supervisor of 2 PhD students; A Bourne and Varsha Pathare. He also supervisors postdoctoral fellows.

Areas of Research / Teaching Expertise

Tree Nutrition and Physiology; Global Change Biology; Ecological Physiology; Plant Physiology; Ecosystem Adaptation to Climate Change; Ecosystem Adaptation in Australia; Ecosystem Adaptation in the World; Forest and Woodland Flora, Fauna and Biodiversity

Awards and Recognition

  • Appointed to the Australian Research Council (ARC) College of Experts, reflecting his enormous contribution to the biological sciences and positioning him as a leading expert in science
  • Appointed to the prestigious ARC Medical Research Advisory Group

^ Back to top

Grants / Current Projects

Quantifying the recovery trajectory and aboveground productivity in rainforest species after fire on rich versus poor soils
Co-Researchers: Kristine Crous, Brendan Choat, Rachael Nolan, and Robert Kooyman (MQ)
Partner/Funding Body: Herman Slade
Period: 2022

Mechanistic responses of phosphorus-limited forests to CO2 enrichment
Co-Researchers: Ben Smith and Catriona Macdonald
Partner/Funding Body: Australian Research Council Discovery
Period: 2022-2025

Feast or famine: how Australian plants stay productive under low phosphorus
Partner/Funding Body: Australian Research Council Discovery
Period: 2021-2023

Curating museum collections for climate change mitigation
Co-Researcher: Fiona Cameron
Partner/Funding Body: Australian Research Council Linkage
Period: 2021-2023

Phosphorus cycle interactions with elevated carbon dioxide in mature forest ecosystems: stand to globe
Co-Researcher: Ben Smith, Catriona Macdonald
Partner/Funding Body: Swedish Research Council [via Lund University]
Period: 2020

Heat Testing: Anzac Parade Planter Bed for Hebe
Partner/Funding Body: National Capital Authority
Period: 2020

Terrestrial Ecosystems Research Network (TERN)Co-Researcher: Matthias Boer, Elise Pendall and Brendan Choat
Partner/Funding Body: NSW Department of Planning, Industry and Environment RAAP
Period: 2019

New paradigms for understanding ecosystem nutrient limitation
Partner/Funding Body: New Phytologist Trust
Period: 2019

Improvement and management of teak and sandalwood in PNG and Australia
Co-Researcher: Tony Page and Braden Jenkin
Partner/Funding Body: Australian Centre for International Agricultural Research
Period: 2019

Terrestrial Ecosystems Research Network – Cumberland Plain Supersite.
Co-Researcher: Matthias Boer, Elise Pendall and Brendan Choat
Partner/Funding Body: NSW Government, Department of Industry, Research Attraction and Acceleration Program (RAAP)
Period: 2018-2022

Data-model integration for understanding nitrogen and phosphorus limitations to future forests
Co-Researchers: Mingkai Jiang
Partner/Funding Body: Australia-Germany Joint Research Co-operation (DAAD) [via the Australian Academy of Sciences]
Period: 2017-2018

Will trees get enough nitrogen to sustain productivity in elevated CO2?
Co-Researchers: Yolima Carrillo, Oula Ghannoum and Sönke Zaehle
Partner/Funding Body: Australian Research Council Discovery
Period: 2016-2018

Which plant where, when and why database for growing urban green space
Co-Researchers: Ian Anderson, Sally Power, Mark Tjoelker and Paul Rymer
Partner/funding body: Horticulture Australia Innovation and NSW Office of Environment and Heritage
Period: 2016-2021

Carbon Water Linkages for native trees under elevated CO2: A unique coupled sensor system for tree water-use efficiency
Co-Researcher: Brendan Choat
Partner/Funding Body: Campbell Scientific Australia Pty Ltd
Period: 2014

When fire and water mix: do carbon dioxide-related water savings drive woody plant thickening and fire dynamics in a grassy woodland?Co-Researchers: Matthias Boer, Victor Resco de Dios and Ross Bradstock
Partner/Funding Body: Australian Research Council Discovery
Period: 2013-2015

What will be the hydrological consequences of elevated CO2? An experimental test of the impacts of elevated CO2 on evapotranspiration in a mature Australian Woodland
Co-researcher: David Tissue
Partner/Funding Body: Commonwealth Scientific and Industrial Research Organisation
Period: 2011-12

Woodland response to elevated CO2 in free air carbon dioxide enrichment: Does phosphorus limit the sink for Carbon?Co-researcher: David Tissue
Partners/Funding Body: Australian Research Council
Period: 2011-2015

Insect herbivore and plant responses in eucalypt forests under climate change at physiological, species and community scales
Co-Researcher: Markus Riegler
Partner/Funding Body: Australian Research Council Discovery
Period: 2010-2012

Forest ecosystem water use: Does species diversity matter?
Co-Researchers: Belinda Medlyn, Charles Morris and Danelia Drewry
Partner/Funding Body: Australian Research Council Linkage
Period: 2009-2011

Understanding the impact of climate change on carbon metabolism in Eucalyptus trees
Co-Researcher: David Tissue
Partner/Funding Body: University of Western Sydney
Period: 2009-2010

How will Eucalypt tree architecture and growth adapt to future atmospheric CO2 and drought?Co-Researchers: Belinda Medlyn, Gabriel Katul and Reinhardt Ceulemans
Partner/Funding Body: Australian Research Council Discovery
Period: 2008-2010

Tunable laser diode for laboratory and field measurements
Co-Researchers: Charles Warren, Mark Adams, Jann Conroy and David Tissue
Partner/Funding Body: Australian Research Council Linkage Infrastructure, Equipment & Facilities
Period: 2008

Trade Off between carbon sequestration in forests and water yields under rising CO2
Co-Researchers: Jann Conroy and David Tissue
Partner/Funding Body: NSW Department of Environment and Climate Change
Period: 2007–2009

The Hawkesbury Forest Experiment: Impacts of Precipitation and CO2 on Trees
Co-Researchers: Jann Conroy, David Tissue, Ross McMurtrie, Derek Eamus, Anthony Haigh, Belinda Medlyn and Sune Linder
Partner/Funding Body: Australian Greenhouse Office
Period: 2005-2008

The Duke Forest FACE Experiment
Partner/Funding Body: U.S. DOE, Office of Science, Biological and Environmental Research
Period: 2001-2004 and 2004-2006

Biodiversity, carbon dioxide (CO2) and Nitrogen: Integration and interactions among plants, consumers, decomposers and mutualists
Co-Researchers: Peter Reich and D Zak
Partner/Funding Body: NSF Biocomplexity
Period: 2003-2007

Canopy conductance and CO2 uptake in response to trace gases CO2 and ozone in aspen and aspen-birch forests
Co-Researchers: M Kubiske and E MacDonald
Partner/Funding Body: U.S. DOE, National Institute for Global Environmental Change, Midwestern Regional Center
Period: 2003-2005

Plant species economics and strategy-dimensions of plant ecology
Co-Researcher: Mark Westoby
Partner/Funding Body: Australian Research Council Discovery
Period: 2003-2005

Northern Michigan forest productivity across a complex landscape
Co-Researcher: K Bergen
Partner/Funding Body: U.S.D.A. McIntire-Stennis Research Program
Period: 2002-2004

^ Back to top

Selected Publications

Cameron FR, Dibley B, Ellsworth DS, (2023) 'Climate Collections and Photosynthetic, Fossil-Fueled Atmospheres', Environmental Humanities, vol.15, no.2, pp 62-84

Crous KY, Cheesman AW, Middleby K, Rogers EIE, Wujeska-Klause A, Bouet AYM, Ellsworth DS, Liddell MJ, Cernusak LA, Barton CVM, (2023) 'Similar patterns of leaf temperatures and thermal acclimation to warming in temperate and tropical tree canopies', Tree Physiology, vol.43, no.8, pp 1383 - 1399

Gardner A, Jiang MK, Ellsworth DS, MacKenzie AR, Pritchard J, Bader MKF, Barton CVM, Bernacchi C, Calfapietra C, Crous KY, Dusenge ME, Gimeno TE, Hall M, Lamba S, Leuzinger S, Uddling J, Warren J, Wallin G, Medlyn BE, (2023) 'Optimal stomatal theory predicts CO2 responses of stomatal conductance in both gymnosperm and angiosperm trees', New Phytologist, vol.237, no.4, pp 1229-1241

Griebel A, Boer MM, Blackman C, Choat B, Ellsworth DS, Madden P, Medlyn B, Resco de Dios V, Wujeska-Klause A, Yebra M, Younes Cardenas N, Nolan RH, (2023) 'Specific leaf area and vapour pressure deficit control live fuel moisture content', Functional Ecology, vol.37, no.3, pp 719 - 731

Liang X, Wang D, Ye Q, Zhang J, Liu M, Liu H, Yu K, Wang Y, Hou E, Zhong B, Xu L, Lv T, Peng S, Lu H, Sicard P, Anav A, Ellsworth DS, (2023) 'Stomatal responses of terrestrial plants to global change', Nature Communications, vol.14, no. 1, Article no.2188

Marchin RM, Medlyn BE, Tjoelker MG, Ellsworth DS, (2023) 'Decoupling between stomatal conductance and photosynthesis occurs under extreme heat in broadleaf tree species regardless of water access', Global Change Biology, vol.29, no.22, pp 6319-6335

Pihlblad J, Andresen LC, Macdonald CA, Ellsworth DS, Carrillo Y, (2023) 'The influence of elevated CO2 and soil depth on rhizosphere activity and nutrient availability in a mature Eucalyptus woodland', Biogeosciences, vol.20, no.3, pp 505 - 521

Zhang Y-B, Corrêa Scalon M, Liu J-X, Song X-Y, Yang D, Zhang Y-J, Ellsworth DS, Zhang J-L, (2023) 'You are what you eat: nutrient and water relations between mistletoes and hosts', New Phytologist, vol.238, no.2, pp 567 - 583

Ellsworth DS, Crous KY, De Kauwe MG, Verryckt LT, Goll D, Zaehle S, Bloomfield KJ, Ciais P, Cernusak LA, Domingues TF, Dusenge ME, Garcia S, Guerrieri R, Ishida FY, Janssens IA, Kenzo T, Ichie T, Medlyn BE, Meir P, Norby RJ, Reich PB, Rowland L, Santiago LS, Sun Y, Uddling J, Walker AP, Weerasinghe KWLK, van de Weg MJ, Zhang Y-B, Zhang J-L, Wright IJ, (2022) 'Convergence in phosphorus constraints to photosynthesis in forests around the world', Nature Communications, vol.13, no.1, Article no.5005

Farrell C, Livesley SJ, Arndt SK, Beaumont L, Burley H, Ellsworth D, Esperon-Rodriguez M, Fletcher TD, Gallagher R, Ossola A, Power SA, Marchin R, Rayner JP, Rymer PD, Staas L, Szota C, Williams NSG, Leishman M, (2022) 'Can we integrate ecological approaches to improve plant selection for green infrastructure?', Urban Forestry and Urban Greening, vol.76, Article no.127732

Gardner A, Ellsworth DS, Crous KY, Pritchard J, MacKenzie AR, (2022) 'Is photosynthetic enhancement sustained through three years of elevated CO2 exposure in 175-year-old Quercus robur?', Tree Physiology, vol.42, no.1, pp 130-144

Gardner A, Ellsworth DS, Pritchard J, MacKenzie AR, (2022) 'Are chlorophyll concentrations and nitrogen across the vertical canopy profile affected by elevated CO2 in mature Quercus trees?', Trees - Structure and Functions, vol.36, no.6, pp 1797-1809

Marchin RM, Backes D, Ossola A, Leishman MR, Tjoelker MG, Ellsworth DS, (2022) 'Extreme heat increases stomatal conductance and drought-induced mortality risk in vulnerable plant species', Global Change Biology, vol.28, no.3, pp 1133-1146

Marchin RM, Esperon-Rodriguez M, Tjoelker MG, Ellsworth DS, (2022) 'Crown dieback and mortality of urban trees linked to heatwaves during extreme drought', Science of the Total Environment, vol.850, Article no.157915

Sabot MEB, De Kauwe MG, Pitman AJ, Ellsworth DS, Medlyn BE, Caldararu S, Zaehle S, Crous KY, Gimeno TE, Wujeska-Klause A, Mu M, Yang J, (2022) 'Predicting resilience through the lens of competing adjustments to vegetation function', Plant Cell and Environment, vol.45, no.9, pp 2744-2761

Sabot MEB, De Kauwe MG, Pitman AJ, Medlyn BE, Ellsworth DS, Martin-StPaul NK, Wu J, Choat B, Limousin J-M, Mitchell PJ, Rogers A, Serbin SP, (2022) 'One Stomatal Model to Rule Them All? Toward Improved Representation of Carbon and Water Exchange in Global Models', Journal of Advances in Modeling Earth Systems, vol.14, no.4, Article no.e2021MS002761

Crous KY, Campany C, Lopez R, Cano F, Ellsworth DS, (2021) 'Canopy position affects photosynthesis and anatomy in mature Eucalyptus trees in elevated CO2', Tree Physiology, vol.41, no.2, pp 206-222

El-Madany TS, Reichstein M, Carrara A, Martin MP, Moreno G, Gonzalez-Cascon R, Penuelas J, Ellsworth DS, Burchard-Levine V, Hammer TW, Knauer J, Kolle O, Luo Y, Pacheco-Labrador J, Nelson A, Perez-Priego O, Rolo V, Wutzler T, Migliavacca,M, (2021) 'How Nitrogen and Phosphorus Availability Change Water Use Efficiency in a Mediterranean Savanna Ecosystem', Journal of Geophysical Research: Biogeosciences, vol.126, no.5, Article no.e2020JG006005

Ely KS, Rogers A, Agarwal DA, Ainsworth EA, Albert LP, Ali A, Anderson J, Aspinwall MJ, Bellasio C, Bernacchi C, Bonnage S, Buckley TN, Bunce J, Burnett AC, Busch FA, Cavanagh A, Cernusak LA, Crystal-Ornelas R, Damerow J, Davidson KJ, De Kauwe MG, Dietze MC, Domingues TF, Dusenge ME, Ellsworth DS, Evans JR, Gauthier PPG, Gimenez BO, Gordon EP, Gough CM, Halbritter AH, Hanson DT, Heskel M, Hogan JA, Hupp JR, Jardine K, Kattge J, Keenan T, Kromdijk J, Kumarathunge DP, Lamour J, Leakey ADB, LeBauer DS, Li Q, Lundgren MR, McDowell N, Meacham-Hensold K, Medlyn BE, Moore DJP, Negrón-Juárez R, Niinemets Ü, Osborne CP, Pivovaroff AL, Poorter H, Reed SC, Ryu Y, Sanz-Saez A, Schmiege SC, Serbin SP, Sharkey TD, Slot M, Smith NG, Sonawane BV, South PF, Souza DC, Stinziano JR, Stuart-Haëntjens E, Taylor SH, Tejera MD, Uddling J, Vandvik V, Varadharajan C, Walker AP, Walker BJ, Warren JM, Way DA, Wolfe BT, Wu J, Wullschleger SD, Xu C, Yan Z, Yang D, (2021) 'A reporting format for leaf-level gas exchange data and metadata', Ecological Informatics, vol.61, Article no.101232

Macdonald CA, Anderson IC, Khachane A, Singh BP, Barton CVM, Duursma RA, Ellsworth DS, Singh BK, (2021) 'Plant productivity is a key driver of soil respiration response to climate change in a nutrient-limited soil', Basic and Applied Ecology, vol.50, pp 155-168

Mao Q, Chen H, Gurmesa GA, Gundersen P, Ellsworth DS, Gilliam FS, Wang C, Zhu F, Ye Q, Mo J, Lu X, (2021) 'Negative effects of long-term phosphorus additions on understory plants in a primary tropical forest', Science of the Total Environment, vol.7981, Article no.149306

Mu MY, De Kauwe MG, Ukkola AM, Pitman AJ, Gimeno TE, Medlyn BE, Or D, Yang JY, Ellsworth DS, (2021) 'Evaluating a land surface model at a water-limited site: implications for land surface contributions to droughts and heatwaves', Hydrology and Earth System Sciences, vol.25, no.1, pp 447-471

Noh NJ, Crous KY, Salomon RL, Li JQ, Ellsworth DS, Barton CVM, Pendall E, Tjoelker MG, (2021) 'Elevated CO2 alters the temperature sensitivity of stem CO2 efflux in a mature eucalypt woodland', Environmental and Experimental Botany, vol.188, Article no.104508

Tabassum S, Ossola A, Marchin RM, Ellsworth DS, Leishman MR, (2021) 'Assessing the relationship between trait-based and horticultural classifications of plant responses to drought', Urban Forestry and Urban Greening, vol.61, Article no.127109

Walker AP, De Kauwe MG, Bastos A, Belmecheri S, Georgiou K, Keeling RF, McMahon SM, Medlyn BE, Moore DJP, Norby RJ, Zaehle S, Anderson-Teixeira KJ, Battipaglia G, Brienen RJW, Cabugao KG, Cailleret M, Campbell E, Canadell JG, Ciais P, Craig ME, Ellsworth DS, Farquhar GD, Fatichi S, Fisher JB, Frank DC, Graven H, Gu LH, Haverd V, Heilman K, Heimann M, Hungate BA, Iversen CM, Joos F, Jiang MK, Keenan TF, Knauer J, Korner C, Leshyk VO, Leuzinger S, Liu Y, MacBean N, Malhi Y, McVicar TR, Penuelas J, Pongratz J, Powell AS, Riutta T, Sabot MEB, Schleucher J, Sitch S, Smith WK, Sulman B, Taylor B, Terrer C, Torn MS, Treseder KK, Trugman AT, Trumbore SE, van Mantgem PJ, Voelker SL, Whelan ME, Zuidema PA, (2021) 'Integrating the evidence for a terrestrial carbon sink caused by increasing atmospheric CO2', New Phytologist, vol.229, no.5, pp 2413-2445

Zhang KY, Yang D, Zhang YB, Ellsworth DS, Xu K, Zhang YP, Chen YJ, He FL, Zhang JL, (2021) 'Differentiation in stem and leaf traits among sympatric lianas, scandent shrubs and trees in a subalpine cold temperate forest', Tree Physiology, vol.41, no.11, pp 1992-2003

Castañeda-Gómez L, Walker JKM, Powell JR, Ellsworth DS, Pendall E, Carrillo Y, (2020) 'Impacts of elevated carbon dioxide on carbon gains and losses from soil and associated microbes in a Eucalyptus woodland', Soil Biology and Biochemistry, vol.143, Article no.107734

Crous KY, Ellsworth DS, (2020) 'Probing the inner sanctum of leaf phosphorus: measuring the fractions of leaf P', Plant and Soil, vol.454, no.s 1-2, pp 77-85

Jiang M, Medlyn BE, Drake JE, Duursma RA, Anderson IC, Barton CVM, Boer MM, Carrillo Y, Castaneda-Gomez L, Collins L, Crous KY, De Kauwe MG, dos Santos BM, Emmerson KM, Facey SL, Gherlenda AN, Gimeno TE, Hasegawa S, Johnson SN, Kannaste A, Macdonald CA, Mahmud K, Moore BD, Nazaries L, Neilson EHJ, Nielsen UN, Niinemets U, Noh NJ, Ochoa-Hueso R, Pathare VS, Pendall E, Pihlblad J, Pineiro J, Powell JR, Power SA, Reich PB, Renchon AA, Riegler M, Rinnan R, Rymer PD, Salomon RL, Singh BK, Smith B, Tjoelker MG, Walker JKM, Wujeska-Klause A, Yang JY, Zaehle S, Ellsworth DS, (2020) 'The fate of carbon in a mature forest under carbon dioxide enrichment', Nature, vol.580, no.7802, pp 227-231

Jiang MK, Caldararu S, Zhang HY, Fleischer K, Crous KY, Yang JY, De Kauwe MG, Ellsworth DS, Reich PB, Tissue DT, Zaehle S, Medlyn BE, (2020) 'Low phosphorus supply constrains plant responses to elevated CO2: A meta-analysis', Global Change Biology, vol.26, no.10, pp 5856-5873

Liang XY, Zhang T, Lu XK, Ellsworth DS, BassiriRad H, You CM, Wang D, He PC, Deng Q, Liu H, Mo JM, Ye Q, (2020) 'Global response patterns of plant photosynthesis to nitrogen addition: A meta-analysis', Global Change Biology, vol.26, no.6, pp 3585-3600

Marchin RM, Ossola A, Leishman MR, Ellsworth DS, (2020) 'A Simple Method for Simulating Drought Effects on Plants', Frontiers in Plant Science, vol.10, Article no.1715

Verryckt LT, Ellsworth DS, Vicca S, Van Langenhove L, Penuelas J, Ciais P, Posada JM, Stahl C, Coste S, Courtois EA, Obersteiner M, Chave J, Janssens IA, (2020) 'Can light‐saturated photosynthesis in lowland tropical forests be estimated by one light level?', Biotropica, vol.52, no.6, pp 1183-1193

Verryckt LT, Van Langenhove L, Ciais P, Courtois EA, Vicca S, Penuelas J, Stahl C, Coste S, Ellsworth DS, Posada JM, Obersteiner M, Chave J, Janssens IA, (2020) 'Coping with branch excision when measuring leaf net photosynthetic rates in a lowland tropical forest', Biotropica, vol.52, no.4, pp 608-615

Yang J, Medlyn BE, De Kauwe MG, Duursma RA, Jiang M,  Kumarathunge D, Crous KY, Gimeno TE, Wujeska-Klause A, Ellsworth DS, (2020) 'Low sensitivity of gross primary production to elevated CO2 in a mature eucalypt woodland', Biogeosciences, vol.17, no.2, pp 265-279

Bloomfield KJ, Prentice IC, Cernusak LA, Eamus D, Medlyn BE, Rumman R, Wright IJ, Boer MM, Cale P, Cleverly J, Egerton JJG, Ellsworth DS, Evans BJ, Hayes LS, Hutchinson MF, Liddell MJ, Macfarlane C, Meyer WS, Togashi HF, Wardlaw T, Zhu LL, Atkin OK, (2019) 'The validity of optimal leaf traits modelled on environmental conditions', New Phytologist, vol.221, no.3, pp 1409-1423

Crous KY, Wujeska-Klause A, Jiang MK, Medlyn BE, Ellsworth DS, (2019) 'Nitrogen and Phosphorus Retranslocation of Leaves and Stemwood in a Mature Eucalyptus Forest Exposed to 5 Years of Elevated CO2', Frontiers in Plant Science, vol.10, Article no.664

Jiang MK, Caldararu S, Zaehle S, Ellsworth DS, Medlyn BE, (2019) 'Towards a more physiological representation of vegetation phosphorus processes in land surface models', New Phytologist, vol.222, no.3, pp 1223-1229

Jiang MK, Zaehle S, De Kauwe MG, Walker AP, Caldararu S, Ellsworth DS, Medlyn BE, (2019) 'The quasi-equilibrium framework revisited: analysing long-term CO2 enrichment responses in plant-soil models', Geoscientific Model Development, vol.12, no.5, pp 2069-2089

Kumarathunge DP, Medlyn BE, Drake JE, Tjoelker MG, Aspinwall MJ, Battaglia M, Cano FJ, Carter KR, Cavaleri MA, Cernusak LA, Chambers JQ, Crous KY, De Kauwe MG, Dillaway DN, Dreyer E, Ellsworth DS, Ghannoum O, Han QM, Hikosaka K, Jensen AM, Kelly JWG, Kruger EL, Mercado LM, Onoda Y, Reich PB, Rogers A, Slot M, Smith NG, Tarvainen L, Tissue DT, Togashi HF, Tribuzy ES, Uddling J, Varhammar A, Wallin G, Warren JM, Way DA, (2019) 'Acclimation and adaptation components of the temperature dependence of plant photosynthesis at the global scale', New Phytologist, vol.222, no.2, pp 768-784

Salomon RL, Steppe K, Crous KY, Noh NJ, Ellsworth DS, (2019) 'Elevated CO2 does not affect stem CO2 efflux nor stem respiration in a dry Eucalyptus woodland, but it shifts the vertical gradient in xylem [CO2]', Plant Cell and Environment, vol.42, no.7, pp 2151-2164

Wujeska-Klause A, Crous KY, Ghannoum O, Ellsworth DS, (2019) 'Leaf age and eCO2 both influence photosynthesis by increasing light harvesting in mature Eucalyptus tereticornis at EucFACE', Environmental and Experimental Botany, vol.167, Article no.103857

Wujeska-Klause A, Crous KY, Ghannoum O, Ellsworth DS, (2019) 'Lower photorespiration in elevated CO2 reduces leaf N concentrations in mature Eucalyptus trees in the field', Global Change Biology, vol.25, no.4, pp 1282-1295

Yang J, Duursma RA, De Kauwe MG, Kumarathunge D, Jiang M, Mahmud K, Gimeno TE, Crous KY, Ellsworth DS, Peters J, Choat B, Eamus D, Medlyn BE, (2019) 'Incorporating non-stomatal limitation improves the performance of leaf and canopy models at high vapour pressure deficit', Tree physiology, vol.39, no.12, pp 1961-1974

Bloomfield KJ, Cernusak LA, Eamus D, Ellsworth DS, Prentice IC, Wright IJ, Boer MM, Bradford MG, Cale P, Cleverly J, Egerton JJG, Evans BJ, Hayes LS, Hutchinson MF, Liddell MJ, Macfarlane C, Meyer WS, Prober SM, Togashi HF, Wardlaw T, Zhu LL, Atkin OK, (2018) 'A continental-scale assessment of variability in leaf traits: Within species, across sites and between seasons', Functional Ecology, vol.32, no.6, pp 1492-1506

Cornwell WK, Wright IJ, Turner J, Maire V, Barbour MM, Cernusak LA, Dawson T, Ellsworth D, Farquhar GD, Griffiths H, Keitel C, Knohl A, Reich PB, Williams DG, Bhaskar R, Cornelissen JHC, Richards A, Schmidt S, Valladares F, Korner C, Schulze ED, Buchmann N, Santiago LS, (2018) 'Climate and soils together regulate photosynthetic carbon isotope discrimination within C-3 plants worldwide', Global Ecology and Biogeography, vol.27, no.9, pp 1056-1067

Drake JE, Macdonald CA, Tjoelker MG, Reich PB, Singh BK, Anderson IC, Ellsworth DS, (2018) 'Three years of soil respiration in a mature eucalypt woodland exposed to atmospheric CO2 enrichment', Biogeochemistry, vol.139, no.1, pp 85-101

Gimeno TE, McVicar TR, O'Grady AP, Tissue DT, Ellsworth DS, (2018) 'Elevated CO2 did not affect the hydrological balance of a mature native Eucalyptus woodland', Global Change Biology, vol.24, no.7, pp 3010-3024

Bourne AE, Creek D, Peters JMR, Ellsworth DS, Choat B, (2017) 'Species climate range influences hydraulic and stomatal traits in Eucalyptus species', Annals of Botany, vol.120, no.1, pp 123-133

Ellsworth DS, Anderson IC, Crous KY, Cooke J, Drake JE, Gherlenda AN, Gimeno TE, Macdonald CA, Medlyn BE, Powell JR, Tjoelker MG, Reich PB, (2017) 'Elevated CO2 does not increase eucalypt forest productivity on a low-phosphorus soil', Nature Climate Change, vol.7, no.4, pp 279-282

Facey SL, Fidler DB, Rowe RC, Bromfield LM, Nooten SS, Staley JT, Ellsworth DS, Johnson SN, (2017) 'Atmospheric change causes declines in woodland arthropods and impacts specific trophic groups', Agricultural and Forest Entomology, vol.19, no.1, pp 101-112

Pathare VS, Crous KY, Cooke J, Creek D, Ghannoum O, Ellsworth DS, (2017) 'Water availability affects seasonal CO2-induced photosynthetic enhancement in herbaceous species in a periodically dry woodland', Global Change Biology, vol.23, no.12, pp 5164-5178

Sharwood RE, Crous KY, Whitney SM, Ellsworth DS, Ghannoum O, (2017) 'Linking photosynthesis and leaf N allocation under future elevated CO2 and climate warming in Eucalyptus globulus', Journal of Experimental Botany, vol.68, no.5, pp 1157-1167

De Kauwe MG, Lin YS, Wright IJ, Medlyn BE, Crous KY, Ellsworth DS, Maire V, Prentice IC, Atkin OK, Rogers A, Niinemets U, Serbin SP, Meir P, Uddling J, Togashi HF, Tarvainen L, Weerasinghe LK, Evans BJ, Ishida FY, Domingues TF, (2016) 'A test of the 'one-point method' for estimating maximum carboxylation capacity from field-measured, light-saturated photosynthesis', New Phytologist, vol.210, no.3, pp 1130-1144

Drake JE, Macdonald CA, Tjoelker MG, Crous KY, Gimeno TE, Singh BK, Reich PB, Anderson IC, Ellsworth DS, (2016) 'Short-term carbon cycling responses of a mature eucalypt woodland to gradual stepwise enrichment of atmospheric CO2 concentration', Global Change Biology, vol.22, no.1, pp 380-390

Duursma RA, Gimeno TE, Boer MM, Crous KY, Tjoelker MG, Ellsworth DS, (2016) 'Canopy leaf area of a mature evergreen Eucalyptus woodland does not respond to elevated atmospheric [CO2] but tracks water availability', Global Change Biology, vol.22, no.4, pp 1666-1676

Gimeno TE, Crous KY, Cooke J, O'Grady AP, Osvaldsson A, Medlyn BE, Ellsworth DS, (2016) 'Conserved stomatal behaviour under elevated CO2 and varying water availability in a mature woodland', Functional Ecology, vol.30, no.5, pp 700-709

Medlyn BE, De Kauwe MG, Zaehle S, Walker AP, Duursma RA, Luus K, Mishurov M, Pak B, Smith B, Wang Y-P, Yang X, Crous KY, Drake JE, Gimeno TE, Macdonald CA, Norby RJ, Power SA, Tjoelker MG, Ellsworth DS, (2016) 'Using models to guide field experiments: a priori predictions for the CO2 response of a nutrient- and water-limited native Eucalypt woodland', Global Change Biology, vol.22, no.8, pp 2834-2851

Norby RJ, De Kauwe MG, Domingues TF, Duursma RA, Ellsworth DS, Goll DS, Lapola DM, Luus KA, Mackenzie AR, Medlyn BE, Pavlick R, Rammig A, Smith B, Thomas R, Thonicke K, Walker AP, Yang X, Zaehle S, (2016) 'Model-data synthesis for the next generation of forest free-air CO2 enrichment (FACE) experiments', New Phytologist, vol.209, no.1, pp 17-28

Bourne AE, Haigh AM, Ellsworth DS, (2015) 'Stomatal sensitivity to vapour pressure deficit relates to climate of origin in Eucalyptus species', Tree Physiology, vol.35, no.3, pp 266-278

Crous KY, Osvaldsson A, Ellsworth DS, (2015) 'Is phosphorus limiting in a mature Eucalyptus woodland? Phosphorus fertilisation stimulates stem growth', Plant and Soil, vol.391, no.s 1-2, pp 293-305

Ellsworth DS, Crous KY, Lambers H, Cooke J, (2015) 'Phosphorus recycling in photorespiration maintains high photosynthetic capacity in woody species', Plant, Cell and Environment, vol.38, no.6, pp 1142-1156

Lin Y-S, Medlyn BE, Duursma RA, Prentice IC, Wang H, Baig S, Eamus D, De Dios VR, Mitchell P, Ellsworth DS, De Beeck MO, Wallin G, Uddling J, Tarvainen L, Linderson M-L, Cernusak LA, Nippert JB, Ocheltree TW, Tissue DT, Martin-StPaul NK, Rogers A, Warren JM, De Angelis P, Hikosaka K, Han Q, Onoda Y, Gimeno TE, Barton CVM, Bennie J, Bonal D, Bosc A, Löw M, Macinins-Ng C, Rey A, Rowland L, Setterfield SA, Tausz-Posch S, Zaragoza-Castells J, Broadmeadow MSJ, Drake JE, Freeman M, Ghannoum O, Hutley LB, Kelly JW, Kikuzawa K, Kolari P, Koyama K, Limousin J-M, Meir P, Da Costa ACL, Mikkelsen TN, Salinas N, Sun W, Wingate L, (2015) 'Optimal stomatal behaviour around the world', Nature Climate Change, vol.5, no.5, pp 459-464

Maire V, Wright IJ, Prentice IC, Batjes NH, Bhaskar R, van Bodegom PM, Cornwell WK, Ellsworth D, Niinemets U, Ordonez A, Reich PB, Santiago LS, (2015) 'Global effects of soil and climate on leaf photosynthetic traits and rates', Global Ecology and Biogeography, vol.24, no.6, pp 706-717

Nielsen UN, Prior S, Delroy B, Walker JKM, Ellsworth DS, Powell JR, (2015) 'Response of belowground communities to short-term phosphorus addition in a phosphorus-limited woodland', Plant and Soil, vol.391, no.1-2, pp 321-331

Quentin AG, Crous KY, Barton CVM, Ellsworth DS, (2015) 'Photosynthetic enhancement by elevated CO2 depends on seasonal temperatures for warmed and non-warmed Eucalyptus globulus trees', Tree Physiology, vol.35, no.11, pp 1249-1263

Quentin AG, Pinkard EA, Ryan MG, Tissue DT, Baggett LS, Adams HD, Maillard P, Marchand J, Landhausser SM, Lacointe A, Gibon Y, Anderegg WRL, Asao S, Atkin OK, Bonhomme M, Claye C, Chow PS, Clement-Vidal A, Davies NW, Dickman LT, Dumbur R, Ellsworth DS, Falk K, Galiano L, Grunzweig JM, Hartmann H, Hoch G, Hood S, Jones JE, Koike T, Kuhlmann I, Lloret F, Maestro M, Mansfield SD, Martinez-Vilalta J, Maucourt M, McDowell NG, Moing A, Muller B, Nebauer SG, Niinemets U, Palacio S, Piper F, Raveh E, Richter A, Rolland G, Rosas T, St Joanis B, Sala A, Smith RA, Sterck F, Stinziano JR, Tobias M, Unda F, Watanabe M, Way DA, Weerasinghe LK, Wild B, Wiley E, Woodruff DR, (2015) 'Non-structural carbohydrates in woody plants compared among laboratories', Tree Physiology, vol.35, no.11, pp 1146-1165

Duursma RA, Barton CVM, Lin Y-S, Medlyn BE, Eamus D, Tissue DT, Ellsworth DS, McMurtrie RE, (2014) 'The peaked response of transpiration rate to vapour pressure deficit in field conditions can be explained by the temperature optimum of photosynthesis', Agricultural and Forest Meteorology, vol.189-190, pp 2-10

Facey SL, Ellsworth DS, Staley JT, Wright D, Johnson SN, (2014) 'Upsetting the order: how climate and atmospheric change affects herbivore-enemy interactions', Current Opinion in Insect Science, vol.5, pp 66-74

Gauthier PPG, Crous KY, Ayub G, Duan H, Weerasinghe LK, Ellsworth DS, Tjoelker MG, Evans JR, Tissue DT, Atkin OK, (2014) 'Drought increases heat tolerance of leaf respiration in Eucalyptus globulus saplings grown under both ambient and elevated atmospheric [CO2] and temperature', Journal of Experimental Botany, vol.65, no.22, pp 6471-6485

Sun Y, Gu L, Dickinson RE, Pallardy SG, Baker J, Cao Y, Damatta FM, Dong X, Ellsworth D, Van Goethem D, Jensen AM, Law BE, Loos R, Martins SCV, Norby RJ, Warren J, Weston D, Winter K, (2014) 'Asymmetrical effects of mesophyll conductance on fundamental photosynthetic parameters and their relationships estimated from leaf gas exchange measurements', Plant, Cell and Environment, vol.37, no.4, pp 978-994

Crous KY, Quentin AG, Lin Y-S, Medlyn BE, Williams DG, Barton CVM, Ellsworth DS, (2013) 'Photosynthesis of temperate Eucalyptus globulus trees outside their native range has limited adjustment to elevated CO2 and climate warming', Global Change Biology, vol.19, no.12, pp 3790-3807

De Kauwe MG, Medlyn BE, Zaehle S, Walker AP, Dietze MC, Hickler T, Jain AK, Luo Y, Parton WJ, Prentice IC, Smith B, Thornton PE, Wang S, Wang Y-P, Wårlind D, Weng E, Crous KY, Ellsworth DS, Hanson PJ, Seok Kim H, Warren JM, Oren R, Norby RJ, (2013) 'Forest water use and water use efficiency at elevated CO2: A model-data intercomparison at two contrasting temperate forest FACE sites', Global Change Biology, vol.19, no.6, pp 1759-1779

Franks PJ, Adams MA, Amthor JS, Barbour MM, Berry JA, Ellsworth DS, Farquhar GD, Ghannoum O, Lloyd J, McDowell N, Norby RJ, Tissue DT, von Caemmerer S, (2013) 'Sensitivity of plants to changing atmospheric CO2 concentration: From the geological past to the next century', New Phytologist, vol.197, no.4, pp 1077-1094

Héroult A, Lin Y-S, Bourne A, Medlyn BE, Ellsworth DS, (2013) 'Optimal stomatal conductance in relation to photosynthesis in climatically contrasting Eucalyptus species under drought', Plant, Cell and Environment, vol.36, no.2, pp 262-274

Lin YS, Medlyn BE, De Kauwe MG, Ellsworth DS, (2013) 'Biochemical photosynthetic responses to temperature: how do interspecific differences compare with seasonal shifts?', Tree Physiology, vol.33, no.8, pp 793-806

Murray TJ, Ellsworth DS, Tissue DT, Riegler M, (2013) 'Interactive direct and plant-mediated effects of elevated atmospheric [CO2] and temperature on a eucalypt-feeding insect herbivore', Global Change Biology, vol.19, no.5, pp 1407-1416

Murray TJ, Tissue DT, Ellsworth DS, Riegler M, (2013) 'Interactive effects of pre-industrial, current and future [CO2] and temperature on an insect herbivore of Eucalyptus', Oecologia, vol.171, no.4, pp 1025-1035

Barton CVM, Duursma RA, Medlyn BE, Ellsworth DS, Eamus D, Tissue DT, Adams MA, Conroy J, Crous KY, Liberloo M, Löw M, Linder S, McMurtrie RE, (2012) 'Effects of elevated atmospheric [CO2] on instantaneous transpiration efficiency at leaf and canopy scales in Eucalyptus saligna', Global Change Biology, vol.18, no.2, pp 585-295

Crous KY, Zaragoza-Castells J, Ellsworth DS, Duursma RA, Löw M, Tissue DT, Atkin OK, (2012) 'Light inhibition of leaf respiration in field-grown Eucalyptus saligna in whole-tree chambers under elevated atmospheric CO2 and summer drought', Plant, Cell and Environment, vol.35, no.5,pp 966-981

Duursma RA, Falster DS, Valladares F, Sterck FJ, Pearcy RW, Lusk CH, Sendall KM, Nordenstahl M, Houter NC, Atwell BJ, Kelly N, Kelly JWG, Liberloo M, Tissue DT, Medlyn BE, Ellsworth DS, (2012) 'Light interception efficiency explained by two simple variables: A test using a diversity of small- to medium-sized woody plants', New Phytologist, vol.193, no.2, pp 397-408

Ellsworth DS, Thomas R, Crous KY, Palmroth S, Ward E, Maier C, Delucia E, Oren R, (2012) 'Elevated CO2 affects photosynthetic responses in canopy pine and subcanopy deciduous trees over 10 years: A synthesis from Duke FACE', Global Change Biology, vol.18, no.1, pp 223-242

Falster DS, Reich PB, Ellsworth DS, Wright IJ, Westoby M, Oleksyn J, Lee TD, (2012) 'Lifetime return on investment increases with leaf lifespan among 10 Australian woodland species', New Phytologist, vol.193, no.2, pp 409-419

Leigh A, Sevanto S, Ball MC, Close JD, Ellsworth DS, Knight CA, Nicotra AB, Vogel S, (2012) 'Do thick leaves avoid thermal damage in critically low wind speeds?', New Phytologist, vol.194, no.2, pp 477-487

Lin YS, Medlyn BE, Ellsworth DS, (2012) 'Temperature responses of leaf net photosynthesis: the role of component processes', Tree Physiology, vol.32, no.2, pp 219-231

Medlyn BE, Duursma RA, Eamus D, Ellsworth DS, Colin Prentice I, Barton CVM, Crous KY, de Angelis P, Freeman M, Wingate L, (2012) 'Corrigendum: Reconciling the optimal and empirical approaches to modelling stomatal conductance', Global Change Biology, vol.18, no.11, p 3476

Crous KY, Zaragoza-Castells J, Löw M, Ellsworth DS, Tissue DT, Tjoelker MG, Barton,CVM, Gimeno TE, Atkin OK, (2011) 'Seasonal acclimation of leaf respiration in Eucalyptus saligna trees: Impacts of elevated atmospheric CO2 and summer drought', Global Change Biology, vol.17, no.4, pp 1560-1576

Dronova I, Bergen KM, Ellsworth DS, (2011) 'Forest Canopy Properties and Variation in Aboveground Net Primary Production over Upper Great Lakes Landscapes', Ecosystems, vol.14, no.6, pp 865-879

Duursma RA, Barton CVM, Eamus D, Medlyn BE, Ellsworth DS, Forster MA, Tissue DT, Linder S, McMurtrie RE, (2011) 'Rooting depth explains [CO2] x drought interaction in Eucalyptus saligna', Tree Physiology, vol.31, no.9, pp 922-931

Medlyn BE, Duursma RA, Eamus D, Ellsworth DS, Prentice IC, Barton CVM, Crous KY, De Angelis P, Freeman M, Wingate L, (2011) 'Reconciling the optimal and empirical approaches to modelling stomatal conductance', Global Change Biology, vol.17, no.6, pp 2134-2144

Zeppel MJB, Lewis JD, Medlyn B, Barton CVM, Duursma RA, Eamus D, Adams MA, Phillips N, Ellsworth DS, Forster MA, Tissue DT, (2011) 'Interactive effects of elevated CO2 and drought on nocturnal water fluxes in Eucalyptus saligna', Tree Physiology, vol.31, no.9, pp 932-944

Barton CVM, Ellsworth DS, Medlyn BE, Duursma RA, Tissue DT, Adams MA, Eamus D, Conroy JP, McMurtrie RE, Parsby J, Linder S, (2010) 'Whole-tree chambers for elevated atmospheric CO2 experimentation and tree scale flux measurements in south-eastern Australia: The Hawkesbury Forest Experiment', Agricultural and Forest Meteorology, vol.150, no.7-8, pp 941-951

Calfapietra C, Ainsworth EA, Beier C, De Angelis P, Ellsworth DS, Godbold DL, Hendrey GR, Hickler T, Hoosbeek M, Karnosky DF, King J, Körner C, Leakey ADB, Lewin KF, Liberloo M, Long SP, Lukac M, Matyssek R, Miglietta F, Nagy J, Norby RJ, Oren R, Percy KE, Rogers A, Scarasci Mugnozza G, Stitt M, Taylor G, Ceulemans R, (2010) 'New challenges and priorities in the next generation of elevated CO2 experiments on forest ecosystems and plantations', Trends in Plant Science, vol.15, no.1, pp 5-10

Crous KY, Reich PB, Hunter MD, Ellsworth DS, (2010) 'Maintenance of leaf N controls the photosynthetic CO2 response of grassland species exposed to 9 years of free-air CO2 enrichment', Global Change Biology, vol.16, no.7, pp 2076-2088

Uddling J, Hogg AJ, Teclaw RM, Carroll MA, Ellsworth DS, (2010) 'Stomatal uptake of O-3 in aspen and aspen-birch forests under free-air CO2 and O-3 enrichment', Environmental Pollution, vol.158, no.6, Special Issue, pp 2023-2031

Cernusak LA, Tcherkez G, Keitel C, Cornwell WK, Santiago LS, Knohl A, Barbour MM, Williams DG, Reich PB, Ellsworth DS, Dawson TE, Griffiths HG, Farquhar GD, Wright IJ, (2009) 'Why are non-photosynthetic tissues generally 13C enriched compared with leaves in C3 plants? Review and synthesis of current hypotheses', Functional Plant Biology, vol.36, no.3, pp 199-213

Franklin O, McMurtrie RE, Iversen CM, Crous KY, Finzi AC, Tissue DT, Ellsworth DS, Oren R, Norby RJ, (2009) 'Forest fine-root production and nitrogen use under elevated CO2: Contrasting responses in evergreen and deciduous trees explained by a common principle', Global Change Biology, vol.15, no.1, pp 132-144

Reich PB, Falster DS, Ellsworth DS, Wright IJ, Westoby M, Oleksyn J, Lee TD, (2009) 'Controls on declining carbon balance with leaf age among 10 woody species in Australian woodland: Do leaves have zero daily net carbon balances when they die?', New Phytologist, vol.183, no.1, pp 153-166

Uddling J, Teclaw RM, Pregitzer KS, Ellsworth DS, (2009) 'Leaf and canopy conductance in aspen and aspen-birch forests under free-air enrichment of carbon dioxide and ozone', Tree Physiology, vol.29, no.11, pp 1367-1380

Crous KY, Walters MB, Ellsworth DS, (2008) 'Elevated CO2 concentration affects leaf photosynthesis-nitrogen relationships in Pinus taeda over nine years in FACE', Tree Physiology, vol.28, no.4, pp 607–614

Uddling J, Teclaw RM, Kubiske ME, Pregitzer KS, Ellsworth DS, (2008) 'Sap flux in pure aspen and mixed aspen-birch forests exposed to elevated concentrations of carbon dioxide and ozone', Tree Physiology, vol.28, no.8, pp 1231-1243

Chung H, Zak DR, Ellsworth DS, Reich PB, (2007) 'Declining plant species richness, elevated CO2, and atmospheric N deposition alter soil microbial community composition and function', Global Change Biology, vol.13, pp 980-989

Chung H, Zak DR, Reich PB, Ellsworth DS, (2007) 'Plant species richness, elevated CO2, and atmospheric nitrogen deposition alter soil microbial community composition and function' Global Change Biology, vol.13, pp 1-10

Hogg AH, Uddling J, Ellsworth DS, Carroll M, Pressley S, Lamb B, Vogel C, (2007) 'Stomatal and non-stomatal fluxes of ozone to a northern mixed hardwood forest', Tellus, vol.59B, pp 514-525

McCarthy HR, Oren R, Finzi AC, Ellsworth DS, Kim H-S, Johnsen KH, Millar B, (2007) 'Temporal dynamics and spatial variability in the enhancement of canopy leaf area under elevated atmospheric CO2', Global Change Biology, vol.13, no.12, pp 2479-2497

Sefcik LT, Zak DR, Ellsworth DS, (2007) 'Seedling survival in a northern temperate forest understory is increased by elevated atmospheric CO2 and atmospheric N deposition', Global Change Biology, vol.13, pp 132-146

Zak DR, Holmes WE, Pregitzer KS, King JS, Ellsworth DS, Kubiske ME, (2007) 'Belowground competition and the response of developing forest communities to atmospheric CO2 and O3', Global Change Biology, vol.13, no.10, pp 2230-2238

Reich PB, Hobbie SE, Lee TD, Ellsworth DS, West JB, Tilman D, Knops JHM, Naeem S, Trost J, (2006) 'Nitrogen limitation constrains sustainability of ecosystem response to CO2', Nature, vol.440, no.7086, pp 922-925

Sefcik LT, Zak DR, Ellsworth DS, (2006) 'Photosynthetic responses to understory shade and elevated CO2 in northern hardwood tree species differing in shade tolerance', Tree Physiology, vol.26, pp 1589-1599

Crous KY, Ellsworth DS, (2004) 'Canopy position affects photosynthetic adjustments to long-term elevated CO2 concentration (FACE) in a mature Pinus taeda L. forest', Tree Physiology, vol.24, pp 961-970

Ellsworth DS, Reich PB, Naumburg ES, Koch GW, Kubiske ME, Smith SD, (2004) 'Photosynthesis, carboxylation and leaf nitrogen responses of 16 species to elevated pCO2 across four free-air CO2 enrichment experiments in forest, grassland and desert', Global Change Biology, vol.10, pp 2121-2138

Novick K, Stoy P, Katul GG, Ellsworth D, Siqueira M, Juang J, Oren R, (2004) 'Carbon dioxide exchange and water vapor in a warm temperate grassland', Oecologia, vol.138, pp 259-274

Nowak RS, Smith SD, Ellsworth DS, (2004) 'Functional responses of plants to elevated atmospheric CO2 – Do photosynthetic and productivity data from FACE experiments support early predictions? Tansley Review', New Phytologist, vol.162, pp 253-280

Reich PB, Naeem S, Tilman D, Ellsworth D, Knops J, Craine J, Trost J, Wedin D, (2004) 'Species and functional group diversity independently influence biomass accumulation and its response to CO2 and N', Proceedings of National Academy of Sciences (USA) , vol.101, pp 10101-10106

Reich PB, Uhl C, Walters M, Prugh L, Ellsworth DS, (2004) 'Births, deaths, longevity and phenology of Amazonian rain forest leaves: results of a 9-year, 40,000 leaf demography census', Ecological Monographs, vol.74, pp 3-24

Eucalyptus Leaf